Iran President Hassan Rouhani. AFP

Iran’s Revolutionary Guards have fired missiles at eastern Syria targeting the ringleaders of the deadly attack on a military parade in Ahvaz.

A statement said “many terrorists” were killed or injured in the strikes.

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Iranian state television suggested that the missiles hit an area close to the border town of Albu Kamal where Islamic State militants are known to operate.

IS and ethnic Arab separatists from Ahvaz both claimed the 22 September parade attack, in which 25 people died.

The Iranian government has alleged that the assailants were jihadist separatists supported by Gulf Arab allies of the United States.

US Defence Secretary Jim Mattis has dismissed the allegation as “ludicrous”.